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The Optimal LAI model predicts seasonal to decadal dynamics of leaf area index based on optimality principles, balancing energy and water constraints.
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This model is based on Zhou et al. (2025), which presents a general model for the seasonal to decadal dynamics of leaf area that combines predictions from both the light use efficiency (LUE) framework and optimization theory.
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## Model Overview
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The optimal LAI model computes LAI dynamically by:
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1. Calculating the seasonal maximum LAI (LAI$_{max}$) based on energy and water limitations
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2. Computing steady-state LAI from daily meteorological conditions
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3. Updating actual LAI using an exponential moving average to represent the lag in leaf development
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## Seasonal Maximum LAI
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The seasonal maximum LAI (LAI$_{max}$) is determined by the minimum of energy-limited and water-limited constraints:
- $A_{0,annual}$ is the annual total potential GPP (mol m⁻² yr⁻¹)
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- $P_{annual}$ is the annual total precipitation (mol m⁻² yr⁻¹)
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- $D_{growing}$ is the mean vapor pressure deficit during the growing season (Pa)
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- $k$ is the light extinction coefficient (dimensionless)
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- $z$ is the unit cost of constructing and maintaining leaves (mol m⁻² yr⁻¹)
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- $c_a$ is the ambient CO₂ partial pressure (Pa)
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- $\chi$ is the ratio of leaf-internal to ambient CO₂ partial pressure (dimensionless)
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- $f_0$ is the fraction of annual precipitation used by plants (dimensionless)
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## Daily Steady-State LAI
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Given daily meteorological conditions, the steady-state LAI ($L_s$) represents the LAI that would be in equilibrium with GPP if conditions were held constant:
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